We don't know-. For instance, it's entirely possible the starting position is just mate in 10000.
However, the obvious answer if we modify the question slightly to the longest forced mate known can be found in an endgame tablebase (a collection of endgame positions, with evaluations and time to mate). The Syzygy tablebase is an example. The current record (ignoring the 50 move rule) is 546 with this position (as mentioned on Wikipedia). Most tablebases ignore the 50 move rule, and as such I cannot find any note on the longest forced mate with the 50 move rule.